title: Variable spatial selection with two stages of migrations and comparisons between different timings.
mesh_terms: D000818:Animals; D004777:Environment; D005787:Gene Frequency; D005808:Genes, Recessive; D005828:Genetics, Population; D005838:Genotype; D006801:Humans; D008954:Models, Biological; D011110:Polymorphism, Genetic; D011157:Population Dynamics; D012641:Selection, Genetic; D012726:Sexual Behavior, Animal