title: Phylogenetic studies of the subgenus Nialoe (s. lat.) (Coleoptera, Carabidae, Genus Pterostichus), part 2: The Asymmetricus species group.
abstract: Pterostichus (Nialoe) asymmetricus Bates and its allies are revised based on the membranous parts of their genitalia. Four new taxa, P. (N.) ovaliphallus sp. nov., P. (N.) fujimurai ibukiyamanus ssp. nov., P. (N.) shotaroi kiimontanus ssp. nov., and P. (N.) basilobatus sp. nov. are described. A cladistic analysis based on 31 morphological characters reveals that Daisenilaoe (s. str.) Nakane and Straneo is included as one clade within Nialoe (s. str.). The speciation and dispersal process of this species group are also discussed.
authors.affiliation: Graduate School of Agricultural and Life Sciences, The University of Tokyo, Yayoi 1-1-1, Bunkyo-ku, Tokyo 113-8657, Japan. ksasa@fr.a.u-tokyo.ac.jp
authors.forename: Kôji
authors.initials: K
authors.lastname: Sasakawa
mesh_terms: D000818:Animals; D001517:Coleoptera / Q000033:anatomy & histology* / Q000145:classification*; D003710:Demography; D005837:Genitalia, Male / Q000033:anatomy & histology*; D005843:Geography; D008297:Male; D010802:Phylogeny; D013045:Species Specificity