title: Tetradactyl footprints of an unknown affinity theropod dinosaur from the Upper Jurassic of Morocco.
abstract: BACKGROUND
  New tetradactyl theropod footprints from Upper Jurassic (Oxfordian-Kimmeridgian) have been found in the Iouaridène syncline (Morocco). The tracksites are at several layers in the intermediate lacustrine unit of Iouaridène Formation. The footprints were named informally in previous works "Eutynichnium atlasipodus". We consider as nomen nudum.
  
  
  RESULTS
  Boutakioutichnium atlasicus ichnogen. et ichnosp. nov. is mainly characterized by the hallux impression. It is long, strong, directed medially or forward, with two digital pads and with the proximal part of the first pad in lateral position. More than 100 footprints in 15 trackways have been studied with these features. The footprints are large, 38-48 cm in length, and 26-31 cm in width.
  
  
  CONCLUSIONS
  Boutakioutichnium mainly differs from other ichnotaxa with hallux impression in lacking metatarsal marks and in not being a very deep footprint. The distinct morphology of the hallux of the Boutakioutichnium trackmaker -i.e. size and hallux position- are unique in the dinosaur autopodial record to date.
authors.affiliation: Faculté des Sciences, Université Mohamed V, Rabat, Morocco.
authors.forename: Jaouad
authors.initials: J
authors.lastname: Nouri
authors.forename: Ignacio
authors.initials: I
authors.lastname: Díaz-Martínez
authors.forename: Félix
authors.initials: F
authors.lastname: Pérez-Lorente
mesh_terms: D000818:Animals; D025061:Dinosaurs / Q000033:anatomy & histology / Q000145:classification; D005580:Fossils; D019015:Geologic Sediments; D009018:Morocco