title: The presence of the recurrent veinlet in the Middle Jurassic Nymphidae (Neuroptera): a unique character condition in Myrmeleontoidea.
abstract: A well-developed recurrent veinlet is found in the forewing of two species of Nymphidae from the Middle Jurassic locality of Daohugou (Inner Mongolia, China), Liminympha makarkini Ren & Engel and Daonymphes bisulca gen. et sp. n. This is the first record of this trait in the clade comprised of the superfamilies Myrmeleontoidea and Chrysopoidea. We interpret the recurrent veinlet in these species as a remnant of the condition present more basally in the psychopsoid + ithonoid + chrysopoid + myrmeleontoid clade (i.e., as a plesiomorphy). Other venational character states of Daonymphes bisulca of interest include the configuration of subcosta anterior (ScA), which is very similar to that of extant Nymphidae. We consider the short ScA terminating on ScP to be an autapomorphy of Neuroptera.
authors.affiliation: College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing, 100048, China ; Institute of Biology and Soil Sciences, Far Eastern Branch of the Russian Academy of Sciences, 100 let Vladivostoku Avenue 159, Vladivostok, 690022, Russia.
authors.forename: Vladimir N
authors.initials: VN
authors.lastname: Makarkin
authors.forename: Qiang
authors.initials: Q
authors.lastname: Yang
authors.forename: Chaofan
authors.initials: C
authors.lastname: Shi
authors.forename: Dong
authors.initials: D
authors.lastname: Ren
keywords: Daohugou; Middle Jurassic; Neuroptera; Nymphidae; recurrent veinlet; subcosta anterior