title: Tales of the unexpected: angiocarpous representatives of the Russulaceae in tropical South East Asia.
abstract: Six new sequestrate Lactarius species are described from tropical forests in South East Asia. Extensive macro- and microscopical descriptions and illustrations of the main anatomical features are provided. Similarities with other sequestrate Russulales and their phylogenetic relationships are discussed. The placement of the species within Lactarius and its subgenera is confirmed by a molecular phylogeny based on ITS, LSU and rpb2 markers. A species key of the new taxa, including five other known angiocarpous species from South East Asia reported to exude milk, is given. The diversity of angiocarpous fungi in tropical areas is considered underestimated and driving evolutionary forces towards gasteromycetization are probably more diverse than generally assumed. The discovery of a large diversity of angiocarpous milkcaps on a rather local tropical scale was unexpected, and especially the fact that in Sri Lanka more angiocarpous than agaricoid Lactarius species are known now.
authors.affiliation: Ghent University, Department of Biology, Research Group Mycology, K.L. Ledeganckstraat 35, 9000 Ghent, Belgium;
authors.forename: A
authors.initials: A
authors.lastname: Verbeken
authors.affiliation: Ghent University, Department of Biology, Research Group Mycology, K.L. Ledeganckstraat 35, 9000 Ghent, Belgium; ; Scientific Institute of Public Health, Section Mycology and Aerobiology, Juliette Wytsmanstraat 14, 1050 Brussels, Belgium.
authors.forename: D
authors.initials: D
authors.lastname: Stubbe
authors.affiliation: Ghent University, Department of Biology, Research Group Mycology, K.L. Ledeganckstraat 35, 9000 Ghent, Belgium;
authors.forename: K
authors.initials: K
authors.lastname: van de Putte
authors.affiliation: Stuttgart State Museum of Natural History, Botany Department, Rosenstein 1, 70191 Stuttgart, Germany.
authors.forename: U
authors.initials: U
authors.lastname: Eberhardt
authors.affiliation: Ghent University, Department of Biology, Research Group Mycology, K.L. Ledeganckstraat 35, 9000 Ghent, Belgium; ; Naturalis Biodiversity Center, P.O. Box 9517, 2300 RA Leiden, The Netherlands.
authors.forename: J
authors.initials: J
authors.lastname: Nuytinck
keywords: Arcangeliella; Lactarius; Martellia; Zelleromyces; gasteroid fungi; hypogeous fungi; morphology; phylogeny