title: Revision of the Late Jurassic teleosaurid genus Machimosaurus (Crocodylomorpha, Thalattosuchia).
abstract: Machimosaurus was a large-bodied genus of teleosaurid crocodylomorph, considered to have been durophagous/chelonivorous, and which frequented coastal marine/estuarine ecosystems during the Late Jurassic. Here, we revise the genus based on previously described specimens and revise the species within this genus. We conclude that there were three European Machimosaurus species and another taxon in Ethiopia. This conclusion is based on numerous lines of evidence: craniomandibular, dental and postcranial morphologies; differences in estimated total body length; geological age; geographical distribution; and hypothetical lifestyle. We re-diagnose the type species Machimosaurus hugii and limit referred specimens to only those from Upper Kimmeridgian-Lower Tithonian of Switzerland, Portugal and Spain. We also re-diagnose Machimosaurus mosae, demonstrate that it is an available name and restrict the species to the uppermost Kimmeridgian-lowermost Tithonian of northeastern France. We re-diagnose and validate the species Machimosaurus nowackianus from Harrar, Ethiopia. Finally, we establish a new species, Machimosaurus buffetauti, for the Lower Kimmeridgian specimens of France and Germany (and possibly England and Poland). We hypothesize that Machimosaurus may have been analogous to the Pliocene-Holocene genus Crocodylus in having one large-bodied taxon suited to traversing marine barriers and additional, geographically limited taxa across its range.
authors.affiliation: School of GeoSciences , University of Edinburgh, The King's Buildings , Edinburgh EH9 3JW, UK ; School of Ocean and Earth Science, National Oceanography Centre , University of Southampton , Southampton SO14 3ZH, UK.
authors.forename: Mark T
authors.initials: MT
authors.lastname: Young
authors.affiliation: Le Musée des Dinosaures d'Espéraza , Espéraza 11260, France.
authors.forename: Stéphane
authors.initials: S
authors.lastname: Hua
authors.affiliation: Department of Earth Sciences , Natural History Museum , London SW7 5BD, UK.
authors.forename: Lorna
authors.initials: L
authors.lastname: Steel
authors.affiliation: School of GeoSciences , University of Edinburgh, The King's Buildings , Edinburgh EH9 3JW, UK ; School of Earth Sciences, Wills Memorial Building , University of Bristol , Bristol BS8 1RJ, UK.
authors.forename: Davide
authors.initials: D
authors.lastname: Foffa
authors.affiliation: School of GeoSciences , University of Edinburgh, The King's Buildings , Edinburgh EH9 3JW, UK ; National Museums Scotland , Chambers Street, Edinburgh , EH1 1JF, UK.
authors.forename: Stephen L
authors.initials: SL
authors.lastname: Brusatte
authors.affiliation: Naturmuseum Solothurn , Klosterplatz 2 , Solothurn 4500, Switzerland.
authors.forename: Silvan
authors.initials: S
authors.lastname: Thüring
authors.affiliation: Departamento de Ciências da Terra, Faculdade de Ciências e Tecnologia , Universidade Nova de Lisboa, GeoBioTec, Quinta da Torre , Caparica 2829-516, Portugal ; Museu da Lourinhã , Rua João Luis de Moura 9 , Lourinhã 2530-158, Portugal.
authors.forename: Octávio
authors.initials: O
authors.lastname: Mateus
authors.affiliation: Museo del Jurásico de Asturias (MUJA) , Colunga 33328, Spain.
authors.forename: José Ignacio
authors.initials: JI
authors.lastname: Ruiz-Omeñaca
authors.affiliation: Paläontologische Sammlung, Senckenberg Center for Human Evolution and Palaeoenvironment , Eberhard Karls Universität, Sigwartstrasse 10 , Tübingen 72072, Germany.
authors.forename: Philipe
authors.initials: P
authors.lastname: Havlik
authors.affiliation: Sciences et Géologie Normandes , Le Havre 76620, France.
authors.forename: Yves
authors.initials: Y
authors.lastname: Lepage
authors.affiliation: Faculdade de Biociências , Pontifícia Universidade Católica do Rio Grande do Sul, Avenida Ipiranga 6681 , Porto Alegre 90619-900, Brazil.
authors.forename: Marco Brandalise
authors.initials: MB
authors.lastname: De Andrade
keywords: Africa; Europe; Kimmeridgian; Machimosaurus; Teleosauridae; Tithonian