title: Fifty Shades of Grey: giving colour to the poorly known Angolan Ashy reed frog (Hyperoliidae: Hyperolius cinereus), with the description of a new species.
abstract: Phylogenetic reconstruction using the mitochondrial 16S marker shows that geographically separated populations of the poorly known Hyperolius cinereus (Anura: Hyperoliidae) from Angola form two distinct clades. The description of H. cinereus was originally based on only a single preserved adult male. Fresh material of both sexes allowed a detailed redescription of the species, which is restricted mainly to the south-draining Cunene and Cubango river systems. Bioacoustic and morphological characters, in conjunction with colouration differences, allow the description of a cryptic sister species from Lagoa Carumbo in north-eastern Angola, occurring in the Luele and Lovuo river systems of the Congo drainage basin. Tadpoles, for H. cinereus and the new species, are described.
authors.affiliation: Port Elizabeth Museum, P.O. Box 13147, Humewood, Port Elizabeth 6013, South Africa. werner@bayworld.co.za
authors.forename: Werner
authors.initials: W
authors.lastname: Conradie
authors.forename: William R
authors.initials: WR
authors.lastname: Branch
authors.forename: Krystal A
authors.initials: KA
authors.lastname: Tolley
mesh_terms: D000810:Angola; D063147:Animal Distribution; D000818:Animals; D017753:Ecosystem; D005260:Female; D007814:Larva; D008297:Male; D010802:Phylogeny; D011898:Ranidae / Q000033:anatomy & histology* / Q000145:classification* / Q000235:genetics / Q000502:physiology; D045483:Rivers; D013045:Species Specificity; D014828:Vocalization, Animal