title: Two new species of Euptychia Hübner, 1818 from the upper Amazon basin (Lepidoptera, Nymphalidae, Satyrinae).
abstract: Two new species of Euptychia Hübner, 1818 are described from the upper Amazon basin: Euptychia attenboroughi Neild, Nakahara, Fratello & Le Crom, sp. n. (type locality: Amazonas, Venezuela), and Euptychia sophiae Zacca, Nakahara, Dolibaina & Dias, sp. n. (type locality: Acre, Brazil). Their unusual facies prompted molecular and phylogenetic analyses of one of the species resulting in support for their classification in monophyletic Euptychia. Diagnostic characters for the two species are presented based on wing morphology, wing pattern, presence of androconial patches on the hindwing, and genitalia. Our results indicate that the projection of the tegumen above the uncus, previously considered a synapomorphy for Euptychia, is not shared by all species in the genus. The adults and their genitalia are documented, and distribution data and a map are provided.
authors.affiliation: Scientific Associate, The Natural History Museum, Life Sciences Dept., Cromwell Rd., London SW7 5BD, UK.
authors.forename: Andrew F E
authors.initials: AF
authors.lastname: Neild
authors.affiliation: McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL 32611, USA.
authors.forename: Shinichi
authors.initials: S
authors.lastname: Nakahara
authors.affiliation: Departamento de Zoologia, Laboratório de Estudos de Lepidoptera Neotropical, Universidade Federal do Paraná - UFPR, Caixa Postal 19020, 81531-980, Curitiba, Paraná, Brasil.
authors.forename: Thamara
authors.initials: T
authors.lastname: Zacca
authors.affiliation: 11 First St., W. Islip, NY 11795, USA.
authors.forename: Steven
authors.initials: S
authors.lastname: Fratello
authors.affiliation: Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Apartado 14-0434, Lima-14, Peru.
authors.forename: Gerardo
authors.initials: G
authors.lastname: Lamas
authors.affiliation: Calle 61, N° 37-31, Bogotá, Colombia.
authors.forename: Jean-François
authors.initials: JF
authors.lastname: Le Crom
authors.affiliation: Departamento de Zoologia, Laboratório de Estudos de Lepidoptera Neotropical, Universidade Federal do Paraná - UFPR, Caixa Postal 19020, 81531-980, Curitiba, Paraná, Brasil.
authors.forename: Diego R
authors.initials: DR
authors.lastname: Dolibaina
authors.affiliation: Departamento de Zoologia, Laboratório de Estudos de Lepidoptera Neotropical, Universidade Federal do Paraná - UFPR, Caixa Postal 19020, 81531-980, Curitiba, Paraná, Brasil.
authors.forename: Fernando M S
authors.initials: FM
authors.lastname: Dias
authors.affiliation: Departamento de Zoologia, Laboratório de Estudos de Lepidoptera Neotropical, Universidade Federal do Paraná - UFPR, Caixa Postal 19020, 81531-980, Curitiba, Paraná, Brasil.
authors.forename: Mirna M
authors.initials: MM
authors.lastname: Casagrande
authors.affiliation: Departamento de Zoologia, Laboratório de Estudos de Lepidoptera Neotropical, Universidade Federal do Paraná - UFPR, Caixa Postal 19020, 81531-980, Curitiba, Paraná, Brasil.
authors.forename: Olaf H H
authors.initials: OH
authors.lastname: Mielke
authors.affiliation: McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL 32611, USA.
authors.forename: Marianne
authors.initials: M
authors.lastname: Espeland
keywords: Brazil; Colombia; Euptychiina; Neotropics; Peru; South America; Venezuela; mitochondrial DNA COI barcode