title: The transfer of two clades of Malaysian Sphenomorphus Fitzinger (Squamata: Scincidae) into the genus Tytthoscincus Linkem, Diesmos, & Brown and the description of a new Malaysian swamp-dwelling species.
abstract: Phylogenetic analyses based on the mitochondrial gene ND2 and its flanking tRNAs indicate the diminutive upland and insular species Sphenomorphus bukitensis, S. butleri, S. langkawiensis, S. perhentianensis, and S. temengorensis form a monophyletic group that is phylogenetically embedded within the Southeast Asian genus Tytthoscincus. The analyses also indicate that a new swamp-dwelling skink from the Bukit Panchor State Park, Pulau Pinang, Peninsular Malaysia is the sister species to the swamp-dwelling species S. sibuensis from Pulau Sibu, Johor and Singapore and that these two are also embedded in the genus Tytthoscincus. By transferring the two Peninsular Malaysian clades of Sphenomorphus into the genus Tytthoscincus, the monophyly of the latter is maintained. The new species T. panchorensis sp. nov. can be distinguished from all other species of Tytthoscincus by having a unique combination of morphological and color pattern characteristics.
authors.affiliation: Herpetology Laboratory, Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92515 USA.; Email: lgrismer@lasierra.edu.
authors.forename: L Lee
authors.initials: LL
authors.lastname: Grismer
authors.affiliation: School of Biological Sciences, Universiti Sains Malaysia, 11800 USM, Pulau Pinang, Penang, Malaysia.; Email: mamuin@gmail.com.
authors.forename: Mohd Abdul
authors.initials: MA
authors.lastname: Muin
authors.affiliation: Department of Biology, Brigham Young University, 150 East Bulldog Boulevard, Provo, Utah 84602 USA.; Email: pwood@byu.edu.
authors.forename: Perry L Jr
authors.initials: PL
authors.lastname: Wood
authors.affiliation: School of Biological Sciences, Universiti Sains Malaysia, 11800 USM, Pulau Pinang, Penang, Malaysia.; Email: shahrulanuar@gmail.com.
authors.forename: Shahrul
authors.initials: S
authors.lastname: Anuar
authors.affiliation: Department of Biology, University of Washington, Seattle, Washington, 98195, USA.; Email: cwlinkem@gmail.com.
authors.forename: Charles W
authors.initials: CW
authors.lastname: Linkem
mesh_terms: D000818:Animals; D002965:Classification; D003710:Demography; D050259:Genes, Mitochondrial; D049810:Genetic Speciation; D008116:Lizards / Q000145:classification* / Q000235:genetics* / Q000502:physiology; D008296:Malaysia; D010802:Phylogeny; D012343:RNA, Transfer / Q000235:genetics*; D053833:Wetlands
chemical_list: D012343:RNA, Transfer