title: A new species of Odorrana (Amphibia: Anura: Ranidae) from Vietnam.
abstract: A new species of Odorrana is described from the karst forests in northeastern Vietnam based on morphological differences and molecular divergence. Morphologically, the new species is distinguishable from its congeners on the basis of a combination of the following diagnostic characters: (1) size large (SVL 85.9-91.6 mm in males, 108.7-110.1 mm in females); (2) head longer than wide; (3) vomerine teeth present; (4) external vocal sacs absent; (5) snout short (SL/SVL 0.16-0.17); (6) tympanum large (TD/ED 0.70 in males, 0.68 in females); (7) dorsal surface of head and anterior part of body smooth, posterior part of body and flanks with small tubercles; (8) supratympanic fold present; (9) dorsolateral fold absent; (10) webbing formula I0-0II0-0III0-1/2IV1/2-0V; (11) in life, dorsum green with dark brown spots; (12) flanks greyish brown with dark brown spots; (13) throat and chest grey, underside of limbs with large dark brown spots, edged in white, forming a network. In the phylogenetic analyses, the new species is unambiguously nested within the O. andersonii group, and placed as the sister taxon to O. wuchuanensis.
authors.affiliation: Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam.; Email: cuong03091983@yahoo.com.
authors.forename: Cuong The
authors.initials: CT
authors.lastname: Pham
authors.affiliation: Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam.; Email: nqt2@yahoo.com.
authors.forename: Truong Quang
authors.initials: TQ
authors.lastname: Nguyen
authors.affiliation: Faculty of Environmental Sciences, Hanoi University of Science, Vietnam National University, 334 Nguyen Trai Road, Hanoi, Vietnam. Centre for Natural Resources and Environmental Studies, Hanoi National University, 19 Le Thanh Tong, Hanoi, Vietnam Department of Herpetology, American Museum of Natural History, Central Park West at 79th Street, New York, New York 10024; Email: le.duc.minh@hus.edu.vn.
authors.forename: Minh Duc
authors.initials: MD
authors.lastname: Le
authors.affiliation: Zoological Institute, Department of Terrestrial Ecology, University of Cologne, Zülpicher Strasse 47b, D-50674 Cologne, Germany.; Email: m.bonkowski@uni-koeln.de.
authors.forename: Michael
authors.initials: M
authors.lastname: Bonkowski
authors.affiliation: Zoological Institute, Department of Terrestrial Ecology, University of Cologne, Zülpicher Strasse 47b, D-50674 Cologne, Germany.; Email: m.bonkowski@uni-koeln.de.
authors.forename: Thomas
authors.initials: T
authors.lastname: Ziegler
mesh_terms: D063147:Animal Distribution; D000825:Animal Structures / Q000033:anatomy & histology / Q000254:growth & development; D000818:Animals; D049628:Body Size; D017753:Ecosystem; D005260:Female; D008297:Male; D009929:Organ Size; D010802:Phylogeny; D011898:Ranidae / Q000033:anatomy & histology / Q000145:classification* / Q000235:genetics / Q000254:growth & development; D014744:Vietnam