title: Additions to Philippine Slender Skinks of the Brachymeles bonitae Complex (Reptilia: Squamata: Scincidae) II: a new species from the northern Philippines.
abstract: We describe a new digitless scincid lizard of the genus Brachymeles from northern Luzon and Camiguin Norte islands in the Philippines. This species belongs to the Brachymeles bonitae Complex, and both molecular and morphological data confirm that this species is distinct from all other congeners. Formerly considered to be a single widespread species, this group of species has been the focus of recent systematic reviews. Here we describe a new species in the B. bonitae Complex, recognized currently to constitute five species. Brachymeles ilocandia sp. nov. is the second digitless and the seventeenth non-pentadactyl species in genus. The description of this species brings the total number of species in the genus to 40, and provides new insight into unique distribution patterns of species of the northern Philippines.
authors.affiliation: Department of Biology, University of Oklahoma, 730 Van Vleet Oval, Norman, OK 73019, USA. Sam Noble Oklahoma Museum of Natural History, University of Oklahoma, 2401 Chautauqua Avenue, Norman, OK 73072, USA.; Email: camsiler@ou.edu.
authors.forename: Cameron D
authors.initials: CD
authors.lastname: Siler
authors.affiliation: Department of Biology, University of South Dakota, 414 East Clark Street, Vermillion, SD 57069, USA.; Email: drew.davis@usd.edu.
authors.forename: Drew R
authors.initials: DR
authors.lastname: Davis
authors.affiliation: Department of Biology, University of Oklahoma, 730 Van Vleet Oval, Norman, OK 73019, USA.; Email: elysefreitas@gmail.com.
authors.forename: Elyse S
authors.initials: ES
authors.lastname: Freitas
authors.affiliation: Department of Biology, University of Oklahoma, 730 Van Vleet Oval, Norman, OK 73019, USA.; Email: nahuron@ou.edu.
authors.forename: Nicholas A
authors.initials: NA
authors.lastname: Huron
authors.affiliation: Department of Biology, University of Oklahoma, 730 Van Vleet Oval, Norman, OK 73019, USA; Email: a.w.amrein@gmail.com.
authors.forename: Aaron D
authors.initials: AD
authors.lastname: Geheber
authors.affiliation: Sam Noble Oklahoma Museum of Natural History, University of Oklahoma, 2401 Chautauqua Avenue, Norman, OK 73072, USA.; Email: jwatters@ou.edu.
authors.forename: Jessa L
authors.initials: JL
authors.lastname: Watters
authors.affiliation: Department of Biology, University of Oklahoma, 730 Van Vleet Oval, Norman, OK 73019, USA; Email: michelle.l.penrod-1@ou.edu.
authors.forename: Michelle L
authors.initials: ML
authors.lastname: Penrod
authors.affiliation: Department of Integrative Biology, Oklahoma State University, 501 Life Sciences West, Stillwater, OK, 74074 USA.; Email: papes@okstate.edu.
authors.forename: Monica
authors.initials: M
authors.lastname: Papeș
authors.affiliation: Department of Biology, University of Oklahoma, 730 Van Vleet Oval, Norman, OK 73019, USA.; Email: annalisa.r.manning-1@ou.edu.
authors.forename: Andrew
authors.initials: A
authors.lastname: Amrein
authors.affiliation: Department of Biology, University of Oklahoma, 730 Van Vleet Oval, Norman, OK 73019, USA.; Email: unknown.
authors.forename: Alyssa
authors.initials: A
authors.lastname: Anwar
authors.affiliation: Department of Biology, University of Oklahoma, 730 Van Vleet Oval, Norman, OK 73019, USA.; Email: unknown.
authors.forename: Dontae
authors.initials: D
authors.lastname: Cooper
authors.affiliation: Department of Biology, University of Oklahoma, 730 Van Vleet Oval, Norman, OK 73019, USA.; Email: unknown.
authors.forename: Tucker
authors.initials: T
authors.lastname: Hein
authors.affiliation: Department of Biology, University of Oklahoma, 730 Van Vleet Oval, Norman, OK 73019, USA.; Email: unknown.
authors.forename: Annalisa
authors.initials: A
authors.lastname: Manning
authors.affiliation: Department of Biology, University of Oklahoma, 730 Van Vleet Oval, Norman, OK 73019, USA.; Email: unknown.
authors.forename: Neeral
authors.initials: N
authors.lastname: Patel
authors.affiliation: Department of Biology, University of Oklahoma, 730 Van Vleet Oval, Norman, OK 73019, USA.; Email: unknown.
authors.forename: Lauren
authors.initials: L
authors.lastname: Pinaroc
authors.affiliation: Herpetology Section, Zoology Division, Philippine National Museum, Rizal Park, Burgos Street, Manila, Philippines.; Email: arvin.diesmos@gmail.com.
authors.forename: Arvin C
authors.initials: AC
authors.lastname: Diesmos
authors.affiliation: University of Santo Tomas, Espana Boulevard, Manila, Philippines.; Email: maediesmos@gmail.com.
authors.forename: Mae L
authors.initials: ML
authors.lastname: Diesmos
authors.affiliation: Department of Biological Sciences, Louisiana State University, 220 Life Sciences Building, Baton Rouge, LA 70803, USA.; Email: carloliveros@gmail.com.
authors.forename: Carl H
authors.initials: CH
authors.lastname: Oliveros
authors.affiliation: Biodiversity Institute and Department of Ecology and Evolutionary Biology, University of Kansas, 1345 Jayhawk Boulevard, Lawrence, KS 66045, USA.; Email: rafe@ku.edu.
authors.forename: Rafe M
authors.initials: RM
authors.lastname: Brown
mesh_terms: D063147:Animal Distribution; D000818:Animals; D005260:Female; D008116:Lizards / Q000033:anatomy & histology* / Q000145:classification* / Q000502:physiology; D008297:Male; D010679:Philippines; D013045:Species Specificity