title: New occurrences of fossilized feathers: systematics and taphonomy of the Santana Formation of the Araripe Basin (Cretaceous), NE, Brazil.
abstract: Here we describe three fossil feathers from the Early Cretaceous Santana Formation of the Araripe Basin, Brazil. Feathers are the most complex multiform vertebrate integuments; they perform different functions, occurring in both avian and non-avian dinosaurs. Despite their rarity, fossil feathers have been found across the world. Most of the Brazilian feather fossil record comes from the Santana Formation. This formation is composed of two members: Crato (lake) and Romualdo (lagoon); both of which are predominantly reduced deposits, precluding bottom dwelling organisms, resulting in exceptional preservation of the fossils. Despite arid and hot conditions during the Cretaceous, life teemed in the adjacency of this paleolake. Feathered non-avian dinosaurs have not yet been described from the Crato Member, even though there are suggestions of their presence in nearby basins. Our description of the three feathers from the Crato laminated limestone reveals that, despite the small sample size, they can be referred to coelurosaurian theropods. Moreover, based on comparisons with extant feather morphotypes they can be identified as one contour feather and two downy feathers. Despite their rareness and low taxonomic potential, fossilized feathers can offer insights about the paleobiology of its owners and the paleoecology of the Araripe Basin.
authors.affiliation: Departamento de Geologia Sedimentar e Ambiental, Universidade de São Paulo, São Paulo, Brasil; Programa de Pós-Graduação em Gequímica e Geotectônica, Instituto de Geociências, Universidade de São Paulo, São Paulo, Brasil.
authors.forename: Gustavo M E M
authors.initials: GM
authors.lastname: Prado
authors.affiliation: Departamento de Geologia Sedimentar e Ambiental, Universidade de São Paulo , São Paulo , Brasil.
authors.forename: Luiz Eduardo
authors.initials: LE
authors.lastname: Anelli
authors.affiliation: Departamento de Geologia Sedimentar e Ambiental, Universidade de São Paulo , São Paulo , Brasil.
authors.forename: Setembrino
authors.initials: S
authors.lastname: Petri
authors.affiliation: Departamento de Geologia Sedimentar e Ambiental, Universidade de São Paulo, São Paulo, Brasil; Programa de Pós-Graduação em Gequímica e Geotectônica, Instituto de Geociências, Universidade de São Paulo, São Paulo, Brasil.
authors.forename: Guilherme Raffaeli
authors.initials: GR
authors.lastname: Romero
keywords: Araripe Basin; Fossil feathers; Systematic paleontology; Taphonomy; Vertebrate paleontology