Molecular and morphological evidence for recognition of two species withinHarpagonella(Amsinckiinae, Boraginaceae).
Recent taxonomic treatments of the genusHarpagonellahave included only one lower taxon,Harpagonella palmeriA. Gray. However, a larger-fruited variety ofHarpagonella palmerifrom Arizona and Sonora was described by I.M. Johnston in 1924. He continued to recognize this taxon - Harpagonella palmeri var. arizonica - in his treatment of the genus in Kearney and Peebles's Arizona Flora in 1960. Here, we provide two lines of molecular evidence and quantitative morphological evidence from calyx characters showing that plants ofHarpagonellafrom Arizona, Sonora, and central Baja California, corresponding to Johnston's var. arizonica, are distinct fromHarpagonella palmeriof southern California and Baja California. We make the new combinationHarpagonella arizonica(I.M. Johnston) Guilliams & B.G. Baldwin,comb. nov.for the plants from Arizona, Sonora, and central Baja California.
Jepson Herbarium and Department of Integrative Biology, University of California Berkeley, 1001 Valley Life Sciences Building #2465, Berkeley CA 94720-2465, United States of America; Santa Barbara Botanic Garden 1212 Mission Canyon Road, Santa Barbara, CA 93105-2126, United States of America.
C Matt
CM
Guilliams
Jepson Herbarium and Department of Integrative Biology, University of California Berkeley, 1001 Valley Life Sciences Building #2465, Berkeley CA 94720-2465, United States of America.
Timothy
T
Jang
Jepson Herbarium and Department of Integrative Biology, University of California Berkeley, 1001 Valley Life Sciences Building #2465, Berkeley CA 94720-2465, United States of America.
Bruce G
BG
Baldwin
Amsinckiinae; Boraginaceae; Harpagonella; Pectocarya