title: A revision of tetrapod footprints from the late Carboniferous of the West Midlands, UK.
abstract: A series of sandstone slabs from Hamstead, Birmingham (West Midlands, UK), preserve an assemblage of tetrapod trackways and individual tracks from the Enville Member of the Salop Formation (late Carboniferous: late Moscovian-Kasimovian). This material has received limited previous study, despite being one of the few British sites to preserve Carboniferous tetrapod footprints. Here, we restudy and revise the taxonomy of this material, and document it using 3D models produced using photogrammetry. The assemblage is dominated by large tracks assigned toLimnopusisp., which were made by early amphibians (temnospondyls). A number of similar but smaller tracks are assigned toBatrachichnus salamandroides(also made by temnospondyls).Dimetropus leisnerianus(made by early synapsids) andDromopus lacertoides(made by lizard-like sauropsids such as araeoscelids) are also present. This ichnofauna contrasts with a slightly stratigraphically older, more extensive and better-studied assemblage from Alveley (Shropshire), which is dominated by small amphibians with relatively rare reptiliomorphs, but which lacksDromopustracks. The presence ofDromopus lacertoidesat Hamstead is consistent with the trend towards increasing aridity through the late Carboniferous. It is possible that the assemblage is the stratigraphically oldest occurrence of this important amniote ichnotaxon.
authors.affiliation: School of Geography, Earth and Environmental Sciences, University of Birmingham , Birmingham , United Kingdom.
authors.forename: Luke E
authors.initials: LE
authors.lastname: Meade
authors.affiliation: School of Geography, Earth and Environmental Sciences, University of Birmingham , Birmingham , United Kingdom.
authors.forename: Andrew S
authors.initials: AS
authors.lastname: Jones
authors.affiliation: School of Geography, Earth and Environmental Sciences, University of Birmingham , Birmingham , United Kingdom.
authors.forename: Richard J
authors.initials: RJ
authors.lastname: Butler
keywords: Batrachichnus; Carboniferous; Dimetropus; Dromopus; Ichnofossil; Limnopus; Synapsid; Temnospondyl; Trackway