title: Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genusTriodontusWestwood.
abstract: The subfamily Orphninae (Coleoptera: Scarabaeidae) is reviewed from Madagascar. A total of four genera and 39 species were found, all being endemic to the island. The following five new species are described: Triodontus ankarafantsikae, Triodontus lemoulti, Triodontus viettei, Triodontus fairmairei, and Triodontus inexpectatus. The following new synonymies are proposed: Orphnus nigrita Brancsik, 1893 is synonym of Triodontus hova (Fairmaire, 1868); Triodontus occidentalis Paulian, 1977 and Orphnus obsoletus Brancsik, 1893 are synonyms of Triodontus nitidulus (Guérin-Méneville, 1844); Triodontus vadoni Paulian, 1977 and Triodontus perrotorum Paulian, 1977 are synonyms of Triodontus owas Westwood, 1852. Lectotypes are designated for the following names: Orphnus nitidulus Guérin-Méneville, 1844 and Orphnidius modestus Benderitter, 1914. Keys, descriptions, illustrations of habitus and male genitalia, and distributional records maps are given for all species.
authors.affiliation: Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, 199034 Sankt-Petersburg, Russia. MECADEV, UMR 7179 MNHN/CNRS, Muséum national d'Histoire naturelle, Entomologie, CP 50, 45 rue Buffon, 75231 Paris cedex 05, France. Universidade Federal de Mato Grosso, Instituto de Biociencias, Departamento de Biologia e Zoologia, Av. Fernando Correa da Costa, 2367, 78060-900 - Cuiaba, MT, Brazil. frolov@scarabaeoidea.com.
authors.forename: Andrey V
authors.initials: AV
authors.lastname: Frolov
authors.forename: Olivier
authors.initials: O
authors.lastname: Montreuil
authors.forename: Lilia A
authors.initials: LA
authors.lastname: Akhmetova
mesh_terms: D063147:Animal Distribution; D000825:Animal Structures / Q000033:anatomy & histology / Q000254:growth & development; D000818:Animals; D049628:Body Size; D001517:Coleoptera / Q000033:anatomy & histology / Q000145:classification* / Q000254:growth & development; D017753:Ecosystem; D005260:Female; D008270:Madagascar; D008297:Male; D009929:Organ Size
keywords: Coleoptera, scarab beetles, Pseudorphnus, Madecorphnus, Renorphnus, Madagascar