title: New species of the earthworm genusZapatadrilus(Clitellata, Acanthodrilidae) from northern Mexico.
abstract: Three new species of the earthworm genus Zapatadrilus are described from Tamaulipas, Mexico: Zapatadrilus aurelius sp. nov., Zapatadrilus huastecus sp. nov., and Zapatadrilus montezumensis sp. nov. Delimitation of the new species was supported by morphological and molecular (genetic distances) evidence. Diagnostic characters of the three new species include: tubulo-racemose prostates in segments 18 and 20, penial setae absent, typhlosole present and intestine beginning in 17/18. Z. montezumensis sp. nov. is separated by its metandric condition, Z. aurelius sp. nov. by the last hearts in 13 and Z. huastecus sp. nov. by the shape of the spermathecae and patterns of genital markings.
authors.affiliation: Laboratorio de Invertebrados del Suelo. Red de Biodiversidad y Sistemática. Instituto de Ecología A. C. Carretera Antigua a Coatepec No. 351. Col. El Haya. Xalapa, Veracruz. C.P. 91070 MEXICO. gabriela.cervantes@posgrado.ecologia.edu.mx.
authors.forename: Gabriela
authors.initials: G
authors.lastname: Cervantes
authors.forename: Carlos
authors.initials: C
authors.lastname: Fragoso
authors.forename: Alejandro Espinosa DE Los
authors.initials: AE
authors.lastname: Monteros
authors.forename: Gerardo
authors.initials: G
authors.lastname: Sánchez-Ramos
authors.forename: Manuel
authors.initials: M
authors.lastname: Lara-Villalón
authors.forename: Manuel DE Jesús
authors.initials: MJ
authors.lastname: Yañez-Pacheco
authors.forename: Jesús Omar
authors.initials: JO
authors.lastname: Lázaro-Castellanos
authors.forename: Samuel W
authors.initials: SW
authors.lastname: James
mesh_terms: D063147:Animal Distribution; D000825:Animal Structures / Q000033:anatomy & histology / Q000254:growth & development; D000818:Animals; D049628:Body Size; D005260:Female; D008297:Male; D008800:Mexico; D009835:Oligochaeta / Q000033:anatomy & histology / Q000145:classification* / Q000254:growth & development; D009929:Organ Size
keywords: Annelida, Benhamiinae, Diplocardiinae, DNAbarcodes, Exxus, Meronephridial, Trigaster, Octochaetidae, Oligochaeta