design: DNA was extracted from 2-4 milligrams of dried grounded lamella with the DNeasy Plant Mini kit (Qiagen, Germantown, USA). The DNA was then fragmented using an M220 Focused-ultrasonicator (Covaris, Woburn, MA, USA) with varied shearing times (3045 seconds) depending on the DNA fragment size profile. The average fragment size in the specimens ranged from 660 to 880 bp for the specimens used for the targeted capture sequencing. Dual-indexed libraries for the targeted capture sequencing were prepared using the NEBNext Ultra II Library Prep kit and the NEBNext Multiplex Oligos for Illumina (Dual Index Primer Set 1), according to the manufacturers protocols (New England BioLabs, Ipswich, MA, USA), although at half the recommended volumes. Dual-indexed genomic libraries were pooled and hybridised with our custom myBaits enrichment panel, following v3.0 manufacturers protocols, with the exception of pooling four or nine libraries per hybridisation reaction and each reaction having a total of  of the recommended volume. Hybridisations were performed at 65C for 20 hours in Vapo Protect Mastercycler 6325 thermocycler (Eppendorf, Arlington, UK). Captured targets were amplified with a KAPA HiFi 2x HotStart ReadyMix PCR kit (Roche, Basel, Switzerland) for 10 cycles, and the PCR products were cleaned using Agencourt AMPure XP magnetic beads. Thirteen enriched libraries were further pooled for sequencing on an Illumina MiSeq platform (Illumina, San Diego, CA, USA) using v2 Nano chemistry (2  250 bp paired-end reads) at the Jodrell Laboratory, Royal Botanic Gardens, Kew.