design: Linked read library was prepared at the Genetic Analysis Services (GAS), University of Otago (Dunedin, New Zealand). DNA from an adult male was extracted using the Nanobind kit and size-selected for fragments over 40kbp using Blue Pippin (Sage Science, USA). A chromium 10X linked read (10x Genomics, USA) library was prepared following the manufacturers instructions. The library was sequenced on the Illumina Nova-seq platform to generate 2x151bp paired-end reads (Garvan Institute, Australia).