design: Mature flies were fed a diet comprising 50% hydrolyzed yeast protein and 50% sucrose for optimal nutrition, with access to water ad libitum in disposable cotton-seep wool boxes (12 cm  6.8 cm  7 cm). Flies were housed in laboratory experimental screen cages (30 cm  30 cm  30 cm) as reported before in (Liu et al., 2018). Each treatment group consisted of 200 adult flies, except for the rearing group. Laboratory conditions were maintained at 271C, 65-70% relative humidity, and a 10L:14D photoperiod. Air quality and humidity were regulated using a column-type air conditioner inverter and an electric floor-mounted water-added air purification humidifier.