design: Genomic DNA was extracted from plant samples (three biological replicates) using the CTAB method. Bisulfite conversion was performed with the EpiArt DNA Enzymatic Methylation Kit, and whole-genome bisulfite sequencing (WGBS) libraries were constructed using the EpiArt DNA Methylation Library Kit for Illumina V (Yeasen Biotechnology, Shanghai, China). Libraries were sequenced on the Illumina NovaSeq platform.