Spores were fixed with formaldehyde according to PhaseGenomics instructions (Seattle, Washington, USA). Hi-C experiments were performed with the Phase Genomics Proximo Hi-C Kit (Fungal). Libraries were prepared usung the Phase Genomics Proximo Hi-C Kit (Fungal) with the four restriction enzymes: DpnII (GATC), DdeI (CTNAG), HinfI (GANTC), and MseI (TTAA).